File: <chalcido.htm>                                        [For educational purposes only]        Glossary            <Principal Natural Enemy Groups >             <Citations>             <Home>

 

Chalcidoidea (Agaonidae, Aphelinidae, Chalcididae, Elasmidae, Encyrtidae, Eucharitidae, Eulophidae, Eupelmidae, Eurytomidae, Leucopsidae, Mymaridae, Mymerommatidae, Ormyridae, Torymidae, Perilampidae, Pteromalidae, Rotoitidae, Signiphoridae, Tanaostigmatidae, Tetracampidae, Torymidae, Trichogrammatidae, etc.) --

  <General References>

 

 

          This is a large Superfamily of important insects that have been deployed successfully in biological control worldwide.  The number of species is debatable but many specialists admit to being aware of only a fraction of those still awaiting discovery.  Their taxonomy is often difficult as the characters used often show wide variation. Most  species are  small or very tiny, some being  less than 0.4 mm in long.  Chalcids occur almost everywhere, but because of their small size they are usually overlooked.. Their habitat is in a variety of situations, principally on flowers and plant foliage. They usually hold their wings flat over the abdomen when at rest, and many seem to jump when they begin to fly.

.

          Wing venation is a key diagnostic, and the antennae are usually elbowed and never contain more than 13 segments.  The pronotum is quadrate and does not reach the tegulae, and there is usually a prepectus present on the side of the thorax.  Most of the chalcids are dark to black in color, but many are blue or green in color with a metallic sheen.  Most species have reduced wing venation, and there is a lot of variation in body shape.  Some have peculiar, even bizarre, shapes. The wings are reduced or absent in some groups.

 

          Most of the chalcids are parasitoids of other insects, attacking the egg or larval stage of their hosts. Most hosts are Lepidoptera, Diptera, Coleoptera, and Homoptera.  Because these host groups contain many agricultural pests, it is obvious that the chalcids are a valuable group for natural control. Many species have been imported into different countries for biological control. The larvae of few species are phytophagous feeding inside seeds, stems, or galls.

 

 

References:   Please refer to  <biology.ref.htm>, [Additional references may be found at: MELVYL Library  ]

 

Annecke, D.P., and R.L. Doutt. 1961. The genera of the Mymaridae (Hymenoptera: Chalcidoidea).  Republic of South Africa, Department of Agricultural Technical Services, Entomology Mem. Vol. 5.  71 p.

 

Alayo D., and L.R. Hernandez. 1978. Introducci6n al estudio de los himen6pteros de Cuba: Superfamilia Chalcidoidea. Academia de Ciencias de Cuba, La Habana. 105 pp

 

 

Argaman, Q. 1990. A synopsis of Perilampus Latreille with descriptions of new genera and species (Hymenoptera: Perilampidae), I. Acta Zoologica Hungarica 36:189--263.

 

Argaman, Q. 1991. A synopsis of Perilampus Latreille with descriptions of new genera and species (Hymenoptera, Perilampidae), II. Acta Zoologica Hungarica 37:1--19.

 

Askew, R.R. 1970. The female of Neodipara masneri Boucek (Hymenoptera: Pteromalidae), a genus and species new to Britain. Entomol.'s Gazette 21:277‑281.

 

Askew, R.R. 1975. The male of Neodipara masneri Boucek (Hymenoptera: Pteromalidae); a correction. Entomol.'s Gazette 26:208‑210.

 

Bendel‑Janssen, B. 1977. Zur Biologie, Okologie und Ethologie der Chalcidoidea (Hym.). Mitteilungen aus der Biologischen Bundesanstalt fur Land‑ und Forstwirtshaft No. 176.163 pp.

 

Boucek, Z. 1958a. Revision der europaischen Tetracampidae (Hym. Chalcidoidea) mit einem Katalog der Arten der Welt. Acta Entomologica Musei Nationalis Pragae 32(491):41‑90.

 

Boucek, Z. 1958b. Eine Cleonyminen‑Studie; Bestimmungsstabelle der Gattungen mit Beschreibungen und Notizen, eingeschlossen einige Eupelmidae (Hym. Chalcidoidea). Acta Entomologica Musei Nationalis Pragae 32(510):353‑386.

 

Boucek, Z. 1970. On some British Megastigmus (Hym. Torymidae), with a revised key to the west European species. Entomol.'s Gazette 21:265‑275.

 

Boucek, Z. 1974a. A revision of the Leucospidae (Hymenoptera: Chalcidoidea) of the world. Bull. of the British Museum (Natural History) (Entomology), Supplement 23. 241 pp.

 

Boucek, Z. 1974b. The pteromalid subfamily Eutrichosomatinae (Hymenoptera: Chalcidoidea). Journal of Entomology (B) 43(2):129‑138.

 

Boucek, Z. 1978a. A generic key to Perilampinae (Hymenoptera, Chalcidoidea), with a revision of Krombeinius n. gen. and Euperilampus Walker. Entomologica Scandinavica 9(4):299‑307.

 

Boucek, Z. 1978b. A study of the non‑podagrionine Torymidae with enlarged hind femora, with a key to the African genera (Hymenoptera). Journal of the Entomol. Soc. of Southern Africa 41(1):91‑134.

 

Boucek, Z. 1980. A revision of the genus Monacon Waterston (Hymenoptera: Chalcidoidea: Perilampinae), parasites of ambrosia beetles (Coleoptera: Platypodidae). Bull. of Entomol. Research 70(1):73‑96.

 

Boucek, Z. 1986. Taxonomic study of chalcidoid wasps (Hymenoptera) associated with gall midges (Diptera: Cecidomyiidae) on mango trees. Bull. of Entomol. Research 76(3):393‑407.

 

Boucek, Z. 1988a. Australasian Chalcidoidea (Hymenoptera): a biosystematic revision of genera of fourteen families, with a reclassification of species. C.A.B. International, Wallingford, England. 832 pp.

 

Boucek, Z. 1988b. An overview of the higher classification of the Chalcidoidea (Parasitic Hymenoptera). Pages 11--23 in Gupta, V.K., ed. Advances in parasitic Hymenoptera research. E.J. Brill, Leiden, The Netherlands. 546 pp.

 

Boucek, Z., and R.R. Askew. 1968a. Index of world Tetracampidae (Hym. Chalcidoidea). Index of entomophagous insects, Part I. V. Delucchi and G. Remaudiere, eds. Le Franc,ois, Paris, France. 19 pp.

 

Boucek, Z., and R.R. Askew. 1968b. Index of Palearctic Eulophidae (excl. Tetrastichinae) (Hym. Chalcidoidea). Index of entomophagous insects. V. Delucchi and G. Remaudiere, eds. Le Franc,ois, Paris, France. 254 pp.

 

Boucek, Z., and J.S. Noyes. 1987. Rotoitidae, a curious new family of Chalcidoidea (Hymenoptera) from New Zealand. Systematic Entomology 12:407‑412.

 

Boucek, Z., and J.‑Y. Rasplus. 1991. Illustrated key to west‑Palearctic genera of Pteromalidae: Hymenoptera‑Chalcidoidea. INRA, Versailles, France. 144 pp.

 

Boucek, Z., A. Watsham, and J.T. Wiebes. 1981. The fig wasp fauna of the receptacles of Ficus thonningii (Hymenoptera, Chalcidoidea). Tijdschrift voor Entomologie 124(5):149‑233.

 

Burks, B.D. 1959. The species of the genus Herbertia Howard (Hymenoptera, Pteromalidae). Proc. of the Entomol. Soc. of Washington 61(6):249‑255.

 

Burks, B.D. 1965. The North Amer. species of Elasmus Westwood (Hymenoptera, Eulophidae). Proc. of the Biological Soc. of Washington 78(25):201‑207.

 

Burks, B.D. 1971a. A synopsis of the genera of the family Eurytomidae (Hymenoptera: Chalcidoidea). Transactions of the Amer. Entomol. Soc. 97(1):1--89.

 

Burks, B.D. 1971b. A North Amer. Elasmus parasitic on Polistes (Hymenoptera: Eulophidae). Journal of the Washington Academy of Science 61(3):194‑196.

 

Clausen, C.P. 1940. The oviposition habits of the Eucharidae (Hymenoptera). Journal of the Washington Academy of Sciences 30(12):504‑516.

 

Clausen, C.P. 1941. The habits of the Eucharidae. Psyche 48(2‑3):57‑69.

 

Clouatre, A., D. Coderre, and D. Gagnon. 1989. Habitat of a new Mymarommatidae found in southern Quebec, Canada (Hymenoptera: Terebrantes). Canad. Entomol. 121:825‑826.

 

Compere, H. 1947. A new genus and species, Eurymyiocnema aphelinoidea (Hymenoptera, Aphelinidae), and a history of the genera Euryischia Riley and Myiocnema Ashmead. Bull. of Entomol. Research 38(3):381‑388.

 

Copland, M.J.W., and P.E. King. 1972. The structure of the female reproductive system in the Torymidae (Hymenoptera: Chalcidoidea). Transactions of the Royal Entomol. Soc. of London 124(2):191--212.

 

Copland, M.J.W., P.E. King, and D.S. Hill. 1973. The structure of the female reproductive system in the Agaonidae (Chalcidoidea, Hymenoptera). Journal of Entomology 48(1):25‑35.

 

Darling, D.C. 1983. A review of the New World species of Euperilampus (Hymenoptera; Chalcidoidea), with notes about host associations and phylogenetic relationships. Quaestiones Entomologicae 19(1--2):1--40.

 

Darling, D.C. 1986. Revision of the New World Chrysolampinae (Hymenoptera: Chalcidoidea). Canad. Entomol. 118(9):913--940.

 

Darling, D.C. 1988. Comparative morphology of the labrum in Hymenoptera: the digitate labrum of Perilampidae and Eucharitidae (Chalcidoidea). Canad. Journal of Zoology 66(12):2811‑2835.

 

Darling, D.C. 1991. Revision of the world species of Spalangiopelta (Hymenoptera: Chalcidoidea:

 

Pteromalidae: Ceinae). Royal Ontario Museum Life Sciences Contribution 155. 43 pp.

 

Darling, D.C., and T.D. Miller. 1991. Life history and larval morphology of Chrysolampus (Hymenoptera: Chalcidoidea: Chrysolampinae) in western North Arnerica. Canad. Journal of Zoology 69:2168‑2177.

 

Debauche, H.R. 1948. Etude sur les Mymarommidae et les Mymaridae de la Belgique (Hymenoptera Chalcidoidea). Memoires du Musee Royal d'Histoire Naturelle de Belgique 108. 248 pp.

 

Delvare, G., and H.‑P. Aberlenc. 1989. Les insectes d'Afrique et d'Amerique tropicale. Cles pour la reconnaissance des familles. PRIFAS, CIRAD‑GERDAT, Montpellier, France. 302 pp.

 

Delvare, G., and Z. Boucek. 1992. On the New World Chalcididae (Hymenoptera). Memoirs of the Amer. Entomol. Institute No. 53. 466 pp.

 

De Santis, L. 1948. Estudio monografico de los afelinidos de la Republica Argentina (Hymenoptera, Chalcidoidea). Revista del Museo de la Plata (nueva serie), Seccion Zoologia 5(32):23‑280.

 

DogVanlar, M. 1991. Systematic positions of some taxa in Ormyridae and descriptions of a new species in Ormyrus from Turkey and a new genus in the family (Hymenoptera, Chalcidoidea). Turkiye Entomologije Dergisi 15:1‑13.

 

Domenichini, G. 1966. Index of Palearctic Tetrastichinae (Hym. Eulophidae). Index of Entomophagous Insects. V. Delucchi and G. Remaudiere, eds. Le Franc,ois, Paris, France. 101 pp.

 

Doutt, R.L., and G. Viggiani. 1968. The classification of the Trichogrammatidae (Hymenoptera: Chalcidoidea). Proc. of the California Academy of Sciences 35(20):477‑586.

 

Ferriere, Ch. 1929. The Asiatic and African species of the genus Elasmus, Westw. (Hym., Chalcid.). Bull. of Entomol. Research 20(4):411‑423.

 

Ferriere, Ch. 1947. Les especes europeennes du genre Elasmus Westw. (Hym. Chalc.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 20(6):565‑580.

 

Ferriere, Ch. 1965. Hymenoptera: Aphelinidae d'Europe et du Bassin Mediterraneen. Masson, Paris, France. 206 pp.

 

Flanders, S.E. 1967. Deviate‑ontogenies in the aphelinid male (Hymenoptera) associated with the ovipositional behaviour of the parental female. Entomophaga 12(5):415‑427.

 

Gahan, A.B. 1946. Review of some chalcidoid genera related to Cerocephala Westwood. Proc. of the United States National Museum 96(3203):349‑376.

 

Gahan, A.B., and Ch. Ferriere. 1947. Notes on some gall‑inhabiting Chalcidoidea (Hymenoptera). Annals of the Entomol. Soc. of Amer. 40(2):271‑302.

 

Gauld, I., and B. Bolton, eds. 1988. The Hymenoptera. Oxford University Press, Oxford, England. xi + 332 pp., 10 color plate.

 

Gibson, G.A.P. 1986a. Evidence for monophyly and relationships of Chalcidoidea, Mymaridae, and Mymarommatidae (Hymenoptera: Terebrantes). Canad. Entomol. 118(3):205‑240.

 

Gibson, G.A.P. 1986b. Mesothoracic skeletomusculature and mechanics of flight and jumping in Eupelminae (Hymenoptera, Chalcidoidea: Eupelmidae). Canad. Entomol. 118(7):691‑728.

 

Gibson, G.A.P. 1989. Phylogeny and classification of Eupelmidae, with a revision of the world genera of Calosotinae and Metapelmatinae (Hymenoptera: Chalcidoidea). Memoirs of the Entomol. Soc. of Canada No. 149. 121 pp.

 

Gordh, G. 1975. The comparative external morphology and systematics of the neotropical parasitic fig wasp genus Idarnes (Hymenoptera: Torymidae). University of Kansas Science Bull. 50(9):389--455.

 

Gordh, G. 1979. Chalcidoidea. Pages 743‑1043 in Krombein, K.V., P.D. Hurd, D.R. Smith, and B.D. Burks, eds. Catalog of Hymenoptera in America north of Mexico, Vol. 1. Smithsonian Institution Press, Washington, D.C., USA. 1198 pp.

 

Graham, M.W.R. de V. 1959. Keys to the British genera and species of Elachertinae, Eulophinae, Entedontinae, and Euderinae (Hym., Chalcidoidea). Transactions of the Soc. for British Entomology 13(10):169‑204.

 

Graham, M.W.R. de V. 1969. The Pteromalidae of northwestern Europe (Hymenoptera: Chalcidoidea). Bull. of the British Museum (Natural History) Entomology, Supplement No. 16. 908 pp.

 

Graham, M.W.R. de V. 1987. A reclassification of the European Tetrastichinae (Hymenoptera:

 

Eulophidae), with a revision of certain genera. Bull. of the British Museum (Natural History), Entomology Series 55(1). 392 pp.

 

Graham, M.W.R. de V. 1991. A reclassification of the European Tetrastichinae (Hymenoptera: Eulophidae): revision of the remaining genera. Memoirs of the Amer. Entomol. Institute 49. 322 pp.

 

Grandi, G. 1929. Studio morfologico e biologico della Blastophaga psenes (L.) (2a edizione riveduta). Bollettino del Laboratorio di Entomologia del R. Istituto Superiore Agrario di Bologna 2:1‑147.

 

Grandi, G. 1963. Catalogo ragionato degli Agaonidi del mondo descritti fino a oggi (6a edizione). Bollettino dell'Istituto di Entomologia della Universita degli studi di Bologna 26:319‑373.

 

Grissell, E.E. 1980. Chalcidoidea. Pages 1‑50 in Syllabus for Parasitic Hymenoptera Training Session 1, 23‑28 June, 1980, College Park, Maryland, USA. Unpublished.

 

Grissell, E.E. 1985. Some nomenclatural changes in the Chalcidoidea (Hymenoptera). Proc. of the Entomol. Soc. of Washington 87(2):350‑355.

 

Grissell, E.E., and C.E. Goodpasture. 1981. A review of Nearctic Podagrionini, with description of sexual behaviour of Podagrion mantis (Hymenoptera: Torymidae). Annals of the Entomol. Soc. of Amer. 74(2):226‑241.

 

Grissell, E.E., and M.E. Schauff. 1990. A handbook of the families of Nearctic Chalcidoidea (Hymenoptera). Entomol. Soc. of Washington, Handbook No. 1. 85 pp.

 

Hayat, M. 1983. The genera of Aphelinidae (Hymenoptera) of the world. Systematic Entomology 8(1):63‑ 102.

 

Hayat, M., and M. Verma. 1980. The aphelinid subfamily Eriaporinae (Hym.: Chalcidoidea). Oriental Insects 14(1):29‑40.

 

Hedqvist, K.‑J. 1961. Notes on Cleonymidae (Hym. Chalcidoidea). I. Entomologisk Tidskrift 82(1 ‑2):91 ‑ 1 10.

 

Hedqvist, K.‑J. 1969a. Notes on Cerocephalini with description of new genera and species (Hymenoptera: Chalcidoidea: Pteromalidae). Proc. of the Entomol. Soc. of Washington 71(3):449‑467.

 

Hedqvist, K.‑J. 1969b. New genera and species of Diparini with notes on the tribe (Hym., Chalcidoidea). Entomologisk Tidskrift 90(3‑4):174‑202.

 

Hedqvist, K.‑J. 1978. A new subfamily and two new genera and species from the New World (Hymenoptera, Chalcidoidea: Pteromalidae). Entomologica Scandinavica 9(2):135--139.

 

Heraty, J.M. 1985. A revision of the Nearctic Eucharitinae (Hymenoptera: Chalcidoidea: Eucharitidae). Proc. of the Entomol. Soc. of Ontario 116:61‑103.

 

Heraty, J.M. 1989. Morphology of the mesosoma of Kapala (Hymenoptera: Eucharitidae) with emphasis on its phylogenetic implications. Canad. Journal of Zoology 67(1):115‑125.

 

Heraty, J.M., and D.C. Darling. 1984. Comparative morphology of the planidial larvae of Eucharitidae and Perilampidae (Hymenoptera: Chalcidoidea). Systematic Entomology 9(3):309‑328.

 

Hill, D.S. 1967a. Figs (Ficus spp.) and fig‑wasps (Chalcidoidea). Journal of Natural History 1(3):413‑434.

 

Hill, D.S. 1967b. Figs (Ficus spp.) of Hong Kong. Hong Kong University Press, Hong Kong. 130 pp., 65 plate.

 

Huber, J.T. 1986. Systematics, biology, and hosts of the Mymaridae and Mymarommatidae (Insecta: Hymenoptera): 1758‑1984. Entomography 4: 185 ‑ 243.

 

Jasnosh, V.A. 1976. Classification of parasitic Hymenoptera of the family Aphelinidae (Chalcidoidea). Entomologicheskoe Obozrenie 55(1):159‑168. [In Russian.] [English translation: Entomol. Review 55(1):114‑120.]

 

Jasnosh, V.A. 1979. Host‑parasite relations in the family Aphelinidae (Hymenoptera, Chalcidoidea). Entomologicheskoe Obozrenie 58(4):751--761. [In Russian.] [English translation: Entomol. Review (1980) 58(4):61‑70.]

 

Jasnosh, V.A. 1983. A review of the aphelinid genera (Hymenoptera, Aphelinidae) of the world. I. A key to the genera. Entomologicheskoe Obozrenie 62(1):157‑171. [In Russian.] [English translation: Entomol. Review 62(1):145--159.]

 

Kalina, V. 1984. New genera and species of Palearctic Eupelmidae (Hymenoptera, Chalcidoidea). Silvaecultura Tropica et Subtropica 10:1‑29.

 

Kamijo, K. 1962. A revision of the species of the Megastigminae occurring in Japan (Hymenoptera: Chalcidoidea) [Taxonomic studies on the Torymidae of Japan, I]. Insecta Matsumurana 25(1):18‑40.

 

Kamijo, K. 1983. A revision of the genus Elatoides Nikol'skaya (Hymenoptera, Pteromalidae), with description of a new species. Kontyu 51(4):573‑581.

 

LaSalle, J. 1987. New World Tanaostigmatidae (Hymenoptera, Chalcidoidea). Contributions of the Amer. Entomol. Institute 23(1). 181 pp.

 

LaSalle, J., and J.S. Noyes. 1985. New family placement for the genus Cynipencyrtus (Hymenoptera: Chalcidoidea: Tanaostigmatidae). Journal of the New York Entomol. Soc. 93(4):1261‑1264.

 

LaSalle, J., and G.I. Stage. 1985. The chalcidoid genus Leptofoenus (Hymenoptera: Pteromalidae). Systematic Entomology 10:285‑298.

 

Medvedev, G.S., editor‑in‑chief. 1978. Keys to the insects of the European part of the USSR. Volume III: Hymenoptera. Part II. Academy of Sciences of the USSR, Institute of Zoology, No. 120, Nauka, Leningrad, USSR. 758 pp. [In Russian.] [English translation: Amerind, New Delhi, India. 1341 pp.]

 

Miller, C.D. 1970. The Nearctic species of Pnigalio and Sympiesis (Hymenoptera: Eulophidae). Memoirs of the Entomol. Soc. of Canada, No. 68.121 pp.

 

Milliron, H.E. 1949. Taxonomic and biological investigations in the genus Megastigmus with particular reference to the taxonomy of the Nearctic species (Hymenoptera: Chalcidoidea; Callimomidae). Amer. Midland Naturalist 41(2):257‑420.

 

Nagarkatti, S., and H. Nagaraja. 1977. Biosystematics of Trichogramma and Trichogrammatoidea species. Annual Review of Entomology 22:157‑ 176.

 

Naumann, I.D. 1991. Hymenoptera (wasps, bees, ants, sawflies). Pages 916‑1000 in CSIRO, ed. The insects of Australia: a textbook for students and research workers, Vol. 2, pp. 543--1137. Melbourne University Press, Carlton, Australia.

 

Nikolskaya, M.N. 1952. The chalcid fauna of the USSR (Chalcidoidea). Keys to the Fauna of the USSR, Zoological Institute of the Academy of Sciences of the USSRj No. 44, Moscow, USSR. 574 pp. [In Russian.] [English translation: Israel Program for Scientific Translations, Jerusalem, Israel, 1963. 593 pp.]

 

Nikolskaya, M.N., and V.A. Jasnosh. 1966. Aphelinidae of the European part of the U.S.S.R. and the Caucasus (Chalcidoidea, Aphelinidae). Akademiia Nauk SSSR, Zoologicheskii Institutom, Izdatel'stvo 'Nauka', Moscow, Russia, No. 91. 294 pp. [In Russian.]

 

Noyes, J.S. 1978. On the numbers of genera and species of Chalcidoidea (Hymenoptera) in the world. Entomolo~ist's Gazette 29 1~

 

Noyes, J.S. 1980. A review of the genera of Neotropical Encyrtidae (Hymenoptera: Chalcidoidea). Bull. of the British Museum (Natural History) (Entomology) 41(3):107‑253.

 

Noyes, J.S. 1988. Encyrtidae (Insecta: Hymenoptera). Fauna of New Zealand No. 13. 188 pp.

 

Noyes, J.S. 1990a. The number of described chalcidoid taxa in the world that are currently regarded as valid. Pages 9‑10 in Chalcid Forum No. 13. 31 pp. Unpublished newsletter.

 

Noyes, J.S. 1990b. Chalcidoid parasitoids. Pages 247‑262 in D. Rosen, ed. World crop pests, Volume 4B. Armored scale insects: their biology, natural enemies and control. Elsevier, Amsterdam, The Netherlands. 688 pp.

 

Noyes, J.S., and M. Hayat. 1984. A review of the genera of Indo‑Pacific Encyrtidae (Hymenoptera: Chalcidoidea). Bull. of the British Museum (Natural History) (Entomology) 48(3):131‑395.

 

Noyes, J.S., and E.W. Valentine. 1989a. Mymaridae (Insecta: Hymenoptera)--introduction, and review of genera. Fauna of New Zealand No. 17. 95 pp.

 

Noyes, J.S., and E.W. Valentine. 1989b. Chalcidoidea (Insecta: Hymenoptera)-- introduction, and review of genera in smaller families. Fauna of New Zealand No. 18. 91 pp.

 

Peck, O., Z. Boucek, and A. Hoffer. 1964. Keys to the Chalcidoidea of Czechoslovakia (Insecta: Hymenoptera). Memoirs of the Entomol. Soc. of Canada No. 34.125 pp.

 

Prinsloo, G.L. 1980. An illustrated guide to the families of African Chalcidoidea (Insecta: Hymenoptera). Republic of South Africa, Department of Agriculture and Fisheries Science Bull. No. 395. 66 pp.

 

Prinsloo, G.L., and D.P. Annecke. 1979. A key to the genera of Encyrtidae from the Ethiopian region, with descriptions of three new genera (Hymenoptera: Chalcidoidea). Journal of the Entomol. Soc. of Southern Africa 42(2):349‑382.

 

Rasnitsyn, A.P., and R. Kulicka. 1990. Hymenopteran insects in Baltic amber with respect to the overall history of the order. Prace Museum Ziemi 41:53‑64.

 

Riek, E.F. 1967. Australian Hymenoptera Chalcidoidea: Family Eulophidae, subfamily Elasminae. Australian Journal of Zoology 15:145‑ 199.

 

Riek, E.F. 1970. Hymenoptera. Pages 867‑ 959 in The insects of Australia. Melbourne University Press, Carlton, Victoria, Australia. 1029 pp.

 

Rosen, D., and P. DeBach. 1979. Species of Aphytis of the world (Hymenoptera: Aphelinidae). Junk, The Hague, The Netherlands. ix + 801 pp.

 

Schauff, M.E. 1984. The Holarctic genera of Mymaridae (Hymenoptera: Chalcidoidea). Memoirs of the Entomol. Soc. of Washington No. 12. 67 pp.

 

Schauff, M.E. 1991. The Holarctic genera of Entedoninae (Hymenoptera: Eulophidae). Contributions of the Amer. Entomol. Institute 26.109 pp.

 

Shafee, S.A. 1975. A new family of Chalcidoidea (Insecta: Hymenoptera). Records of the Zoological Survey of India 68:21--31.

 

Smulyan, M.T. 1936. A revision of the chalcid flies of the genus Perilampus Latreille occurring in America north of Mexico. Proc. of the United States National Museum 83(2990):369‑412.

 

Stage, G.I., and R.R. Snelling. 1986. The subfamilies of Eurytomidae and systematics of the subfamily Heimbrinae (Hymenoptera: Chalcidoidea). Contributions in Science, Natural History Museum of Los Angeles County No. 375.17 pp.

 

Subba Rao, B.R., and M. Hayat, eds. 1985. The Chalcidoidea (Insecta: Hymenoptera) of India and the adjacent countries. Part I: Reviews of families and keys to families and genera. Oriental Insects 19:163‑310 (+310A--0).

 

Sugonyaev, E.S. 1971. A new subfamily of chalcids, Mongolocampinae subfam. n. (Hymenoptera, Chalcidoidea, Tetracampidae) from Mongolia and Kazakhstan. Entomologicheskoe Obozrenie 50(3):664‑675. [In Russian.] [English translation: Entomol. Review 50(3):377‑383.]

 

Szelenyi, G. 1957. The genera of the subfamily Monodontomerinae (Hym. Chalcidoidea). Annales Historico‑Naturales Musei Nationalis Hungarici, series nova 8:381‑388.

 

Tachikawa, T. 1963. Revisional studies on the Encyrtidae of Japan (Hymenoptera: Chalcidoidea). Memoirs of the Ehime University, Section VI (Agriculture) 9(1):1 ‑264.

 

Tachikawa, T. 1981. Hosts of encyrtid genera in the world (Hymenoptera: Chalcidoidea). Memoirs of the College of Agriculture, Ehime University 25(2):85‑ 110.

 

Trjapitzin, V.A. 1971. Review of genera of Palaearctic encyrtids (Hymenoptera, Encyrtidae). Akademiia Nauk SSSR, Paraziticheskie Nasekomye‑entomofagi, Trudy Vsesoyuznogo entomologicheskogo Obshchestva, Izdatel'stvo "Nauka", Leningradskoe Otdelenie, Leningrad 54:68--155. [In Russian.]

 

Trjapitzin, V.A. 1973a. Classification of parasitic Hymenoptera of the family Encyrtidae (Chalcidoidea). Part I. Review of systems of classifications, the subfamily Tetracneminae Howard, 1892. Entomologicheskoe Obozrenie 52(1):163‑175. [In Russian.] [English translation: Entomol. Review 52(1):118‑125.]

 

Trjapitzin, V.A. 1973b. Classification of parasitic Hymenoptera of the family Encyrtidae (Chalcidoidea). Part II. The subfamily Encyrtinae Walker, 1837. Entomologicheskoe Obozrenie 52(2):416--429. [In Russian.] [English translation: Entomol. Review 52(2):287‑295.]

 

Trjapitzin, V.A. 1977. The characteristic features of the morphology of adult encyrtids (Hymenoptera, Chalcidoidea, Encyrtidae) and their systematic significance. Akademiia Nauk SSSR, Paraziticheskie Nasekomye‑entomofagi, Trudy Vsesoyuznogo entomologicheskogo Obshchestva, Izdatel'stvo "Nauka," Leningradskoe Otdelenie, Leningrad 58:145--199. [In Russian.]

 

Trjapitzin, V.A. 1989. Parasitic Hymenoptera of the Family Encyrtidae of the Palaearctics. Akademiia Nauk SSSR, Leningrad, USSR. 487 pp. [In Russian.]

 

Trjapitzin, V.A., and G. Gordh. 1978a. Review of genera of Nearctic Encyrtidae (Hymenoptera, Chalcidoidea). I. Entomologicheskoe Obozrenie 57(2):364--385. [In Russian.] [English translation: Entomol. Review 57:257‑270.]

 

Trjapitzin, V.A., and G. Gordh. 1978b. Review of genera of Nearctic Encyrtidae (Hymenoptera, Chalcidoidea). II. Entomologicheskoe Obozrenie 57(3):636--653. [In Russian.] [English translation: Entomol. Review (1979)57(3):437‑448.]

 

Ulenberg, S.A. 1983. Morphological description of Apocrypta perplexa Coquerel, the type‑species of the genus (fig wasp parasites; Hymenoptera, Chalcidoidea, Toyrmidae).  Proc.

Koninklijke Nederlandse Akademie van Wetenschappen, Series C 86(1):63‑94.

 

Viggiani, G. 1971. Ricerche sugli Hymenoptera Chalcidoidea. XXVIII. Studio morfologico comparativo dell'armatura genitale esterna maschile dei Trichogrammatidae. Bollettino del Laboratorio di Entomologia Agraria "Filippo Silvestri" di Portici 29:181‑222.

 

Viggiani, G. 1988. A preliminary classification of the Mymaridae (Hymenoptera: Chalcidoidea) based on the external male genitalic characters. Bollettino del Laboratorio di Entomologia Agraria "Filippo Silvestri" di Portici 45:141‑148.

 

Wiebes, J.T. 1966a. The structure of the ovipositing organs as a tribal character in the Indo‑Australian sycophagine Toryrnidae (Hymenoptera, Chalcidoidea). Zoologische Mededelingen 41(9):151‑159.

 

Wiebes, J.T. 1966b. Bornean fig wasps from Ficus stupenda Miquel (Hymenoptera, Chalcidoidea). Tijdschrift voor Entomologie 109(7):163‑192.

 

Wiebes, J.T. 1967. Redescription of Sycophaginae from Ceylon and India, with designation of lectotypes, and a world catalogue of the Otitesellini (Hymenoptera, Chalcidoidea, Torymidae). Tijdschrift voor Entomologie 110(13):399‑442.

 

Wiebes, J.T. 1974. Nigeriella, a new genus of West African fig wasps allied to Elisabethiella Grandi (Hymenoptera, Chalcidoidea, Agaonidae). Zoologische Mededelingen 48(5):29‑42.

 

Wiebes, J.T. 1977. A short history of fig wasp research. Gardener's Bull., Singapore 29 2n7‑ ~

 

Wiebes, J.T. 1981. The fig insects of La Reunion (Hymenoptera, Chalcidoidea). Annales de la Societe Entomologique de France (Nouvelle Serie) 17(4):543‑570.

 

Wiebes, J.T. 1982a. Fig wasps (Hymenoptera). Pages 735‑755 in J.L. Gressitt, ed. Biogeography and ecology of New Guinea. Monographiae Biologicae 42. Junk, The Hague, The Netherlands.

 

Wiebes, J.T. 1982b. The phylogeny of the Agaonidae (Hymenoptera, Chalcidoidea). Netherlands Journal of Zoology 32(3):395‑411.

 

Woolley, J.B. 1986. Signiphoridae or Thysanidae? A review of a problem in family‑level nomenclature (Hymenoptera: Chalcidoidea). Bull. of the Entomol. Soc. of Amer. 32(2):91--96.

 

Woolley, J.B. 1988. Phylogeny and classification of the Signiphoridae (Hymenoptera: Chalcidoidea). Systematic Entomology 13:465‑501.

 

Woolley, J.B. 1990. Signiphoridae. Pages 167‑176 in D. Rosen, ed. World crop pests, Volume 4B. Armored scale insects: their biology, natural enemies and control. Elsevier, Amsterdam, The Netherlands. 688 pp.

 

Yoshimoto, C.M. 1975. Cretaceous chalcidoid fossils from Canadian amber. Canad. Entomol. 107(5):499‑528.

 

Yoshimoto, C.M. 1976. Playaspalangia a new genus of Spalangiinae (Hymenoptera, Chalcidoidea: Pteromalidae) from Mexico. Canad. Entomol. 108(5):475‑478.

 

Yoshimoto, C.M. 1977. Revision of the Diparinae (Pteromalidae: Chalcidoidea) from America north of Mexico. Canad. Entomol. 109(7):1035‑1056.

 

Yoshimoto, C.M. 1978. Two new species of Epiclerus from the New World (Hymenoptera: Chalcidoidea, Tetracampidae). Canad. Entomol. 110(11):1207--1211.

 

Yoshimoto, C.M. 1984. The families and subfamilies of Canadian chalcidoid wasps (Hymenoptera: Chalcidoidea). The Insects and Arachnids of Canada, Part 12. Agriculture Canada Publication 1760, Ottawa, Canada. 149 pp.

 

Yoshimoto, C.M. 1990. A review of the genera of New World Mymaridae. (Hymenoptera: Chalcidoidea). Flora and Fauna Handbook No. 7. Sandhill Crane, Gainesville, Florida, USA. 166 pp.

 

Yoshimoto, C.M., M.A. Kozlov, and V.A. Trjapitzin. 1972. A new subfamily of Mymaridae (Hymenoptera, Chalcidoidea). Entomologicheskoe Obozrenie 51:878‑885. [In Russian.] [English translation: Entomol. Review (1989) 51(4):521‑525.]

 

Yousuf, M., and S.A. Shafee. 1986. Checklist of species and bibliography of the world Trichogrammatidae (Hymenoptera). Indian Journal of Systematic Entomology 3(2):29‑82.

 

Zerova, M.D. 1988. The main trends of evolution and the system of chalcids of the family Eurytomidae (Hymenoptera, Chalcidoidea). Entomologicheskoe Obozrenie 67(3):649‑674. [In Russian.] [English translation: Entomol. Review (1989) 68(2):102‑128.