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-22-
 
          Table
1.  Genera of Coleoptera which
predominate in each intertidal zone 
                
in southern California
 
| Rocky headlands and reefs* ---------------------------------- (a) Area of-red algae down        to mean low water           Diaulota       (b) Area of high tides       Thalassotrechus*    
  Diglotta*    
  Bryothinusa*   
  Diaulota*     (also Aegialites.       Amblopusa*, Endeodes       Liparocephalus* and       Ochthebius in
  central       California)                   (c)  Area of spray       Bryothinusa*     Diaulota*     Thalassotrechus         | Sandy beaches  ---------------------------------- Area
  of fresh seaweed       Cafius*     Cicindela     Dyschirius      Thinopinus*   Area
  of decaying seaweed      Aleochara*    
  Anthicus    
  Bledius*    
  Bryobiota*    
  Cafius*    
  Cercyon     
  Diglotta*    
  Emphyastes    
  Hadrotes*     
  Hypocaccus    
  Motschulkium    
  Neopachylopus    
  Phaleria    
  Phycocoetes    
  Pontamalota*    
  Tarphiota*     Thinusa*   Area
  of dry seaweed      Amblyoderus    
  Anthicus     
  Apsena    
  Catorama     Endeodes     Phaleria     Phyconomus | Mudflats  ----------------------------------- Area
  of daily tides      Berosus     Enochrus     Tropisternis    (All in the water)   Area
  of sea-lettuce      Actinidium    
  Bembidion*    
  Carpelimus*     Cicindela     Myllaena*     Ochthebius     Tachys     Thinobius*     Throscinus                   Area of Salicornia       Bledius*     Carpelimus*     Thinobius*         | 
*
Staphyhinidae.
 
| continue their activities in seawater at a reduced rate of metabolism
  (Meyerdirk, 1969). All of the species, which inhabit the reef area, are
  submarine as they are normally submerged in seawater part of the time. One
  species, Diaulota harteri Moore, is found as deep as mean low water
  where it would sometimes be submerged for as long as 20 hours a day. These
  species spend most of their time in crevices in the rocks or in empty
  barnacle shells, etc. They sometimes come out and wander about at low tide.             It is
  questionable if any of the typical species of the sandy beach could be called |